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Valid Names for North America Species
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Valid Stonefly Names for North America

by B.P. Stark, R.W. Baumann, and R.E. DeWalt

Updated as of 3/19/2009

This static list of names is the most complete listing of valid stonefly names for North American species. The list also provides author names for all species and all states and provinces of USA, Canada, and Mexico and some Mesoamerican countries where the species have been reported in peer reviewed literature. To find complete nomenclatural information, literature associated with the order, families, genera, and species, and photographs of selected species please go to http://plecoptera.speciesfile.org.

Click here for a legend to the provincial, state, and Mesoamerican country codes (opens in a new window).

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EUHOLOGNATHA

CAPNIIDAE
    CAPNIINAE

  • Allocapnia
    • aurora Ricker AL, DC, GA, MD, MS, NC, PA, SC, TN, VA
    • brooksi Ross TN
    • cunninghami Ross & Ricker KY, TN
    • curiosa Frison KY, MD, NY, PA, VA, WV
    • forbesi Frison IL, IN, KY, OH, TN, WV
    • frisoni Ross & Ricker KY, NY, OH, PA, TN, VA, WI, WV
    • frumi Kirchner WV
    • fumosa Ross NC, TN, VA
    • granulata (Claassen) AL, AR, DC, IA, IL, IN, KS, KY, LA, MB, MD, MI, MN, MO, MS, NJ, NY, OH, OK, ON, PA, PQ, TN, TX, VA, WI, WV
    • harperi Kirchner PA, VA, WV
    • illinoensis Frison IL, IN, ME, MN, NY, OH, ON, PQ, VA, WI, WV
    • indianae Ricker IN, KY, NY, OH
    • jeanae Ross AR, MO, OK
    • loshada Ricker TN, VA, WV
    • malverna Ross AR, LA, OK, TX
    • maria Hanson CT, MA, MD, ME, NB, NH, NS, NY, PA, PQ, VA, VT, WV
    • menawa Grubbs & Sheldon AL
    • minima (Newport) CT, MA, ME, MI, MN, NB, NF, NH, NS, NY, ON, PQ, VT, WI
    • mohri Ross & Ricker AR, MO, OK
    • muskogee Grubbs & Sheldon AL
    • mystica Frison AL, AR, GA, IL, IN, KY, MO, MS, OH, TN, VA, WV
    • nivicola (Fitch) AL, CT, DC, DE, IL, IN, KY, MA, MD, NB, NC, NJ, NS, NY, OH, PA, PQ, RI, TN, VA, VT, WI, WV
    • ohioensis Ross & Ricker IN, KY, NY, OH, WV
    • oribata Poulton & Stewart AR
    • ozarkana Ross AR
    • pechumani Ross & Ricker NB, NY, PA, PQ
    • peltoides Ross & Ricker AR, OK
    • perplexa Ross & Ricker TN
    • polemistis Ross & Ricker AL, MS
    • pygmaea (Burmeister) CT, DC, IA, KY, MA, MD, ME, MI, MN, MO, NB, ND, NH, NS, NY, OH, ON, PA, PQ, RI, TN, VA, VT, WI, WV
    • recta (Claassen) AL, CT, DC, DE, GA, IL, IN, KY, LA, MA, MD, ME, MS, NC, NH, NS, NY, OH, ON, PA, PQ, SC, TN, VA, VT, WI, WV
    • rickeri Frison AL, AR, DC, DE, GA, IA, IN, KS, KY, MD, MN, MO, MS, NC, NJ, NY, OH, OK, ON, PA, TN, VA, WI, WV
    • sandersoni Ricker AR, MO, OK
    • sano Grubbs AL
    • sequatchie Kondratieff & Kirchner TN
    • simmonsi Kondratieff & Voshell PA, VA
    • smithi Ross & Ricker AL, IL, IN, KY, OH
    • stannardi Ross NC, TN, VA
    • starki Kondratieff & Kirchner LA, MS
    • tennessa Ross & Ricker AL, TN
    • tsalagi Grubbs AL
    • unzickeri Ross & Yamamoto TN
    • virginiana Frison AL, DE, GA, LA, MS, NC, SC, VA
    • vivipara (Claassen) AR, DC, IA, IL, IN, KS, KY, MD, MI, MN, MO, NE, NY, OH, OK, ON, PA, PQ, TN, VA, WI, WV
    • warreni Ross & Yamamoto AR
    • wrayi Ross DC, DE, GA, MD, NC, PA, SC, TN, VA
    • zola Ricker CT, KY, ME, NB, NY, OH, PA, TN, VA, WV
  • Bolshecapnia
    • gregsoni (Ricker) BC
    • maculata (Jewett) CA
    • milami (Nebeker & Gaufin) AB, BC, CO, ID, MT, NM
    • missiona Baumann & Potter MT
    • rogozera (Ricker) BC
    • sasquatchi (Ricker) BC, WA
    • spenceri (Ricker) AB, BC, MT
  • Capnia
    • arapahoe Nelson & Kondratieff CO
    • barberi Claassen CA
    • californica Claassen AZ, CA, MX, NM
    • caryi Baumann & Jacobi AZ, NM
    • cheama Ricker AB, BC, MT, YK
    • coloradensis Claassen BC, CO, ID, MB, MT, NM, SK, WY, YK
    • confusa Claassen AB, AK, BC, CA, CO, ID, MB, MT, NM, SD, SK, UT, WA, WY
    • coyote Nelson & Baumann CA
    • decepta (Banks) AZ, BJ, CO, MX, NM
    • elongata Claassen BC, CA, OR, WA
    • erecta Jewett OR
    • excavata Claassen AK, BC, CA, WA
    • fialai Nelson & Baumann CA
    • giulianii Nelson & Baumann CA
    • glabra Claassen CA, OR
    • gracilaria Claassen AB, BC, BJ, CA, CO, ID, MB, MT, NM, NV, OR, SD, SK, UT, WA, WY
    • hitchcocki Nelson & Baumann CA
    • hornigi Baumann & Sheldon NV
    • inyo Nelson & Baumann CA
    • jewetti Frison OR
    • kersti Nelson OR
    • lacustra Jewett CA, NV
    • licina Jewett OR, WA
    • lineata Hanson CA, ID
    • mariposa Nelson & Baumann CA
    • melia Frison AK, BC, CA, OR, WA
    • mono Nelson & Baumann CA
    • nana Claassen AB, AK, BC, CO, ID, MT, OR, UT, WA, WY
    • nearctica Banks AK, BC, MB, NT, ON
    • nedia Nebeker & Gaufin ID, OR
    • nelsoni Kondratieff & Baumann CO
    • ophiona Nelson & Baumann CA
    • oregona Frison OR
    • palomar Nelson & Baumann CA
    • petila Jewett AB, BC, ID, MT, OR, UT, WY
    • pileata Jewett BC, CA
    • promota Frison BC, CA, OR, WA
    • quadrituberosa Hitchcock CA
    • regilla Nelson & Baumann CA
    • saratoga Nelson & Baumann CA
    • scobina Jewett CA
    • sequoia Nelson & Baumann CA
    • sextuberculata Jewett AB, BC, MT, OR, WA
    • shepardi Nelson & Baumann CA
    • spinulosa Claassen CA
    • teresa Claassen CA
    • tumida Claassen CA, OR
    • uintahi Gaufin CO, ID, NM, NV, UT, WY
    • umpqua Frison CA, OR
    • utahensis Gaufin & Jewett CA, NV, UT
    • valhalla Nelson & Baumann CA
    • ventura Nelson & Baumann CA
    • vernalis (Newport) AB, BC, CO, ID, LB, MB, MI, MN, MT, NB, NM, OH, ON, PQ, SK, UT, WI, WY
    • willametta Jewett OR
    • yosemite Nelson & Baumann CA
    • zukeli Hanson ID
  • Capnura
    • anas Nelson & Baumann OR
    • elevata (Frison) OR, WA
    • fibula (Claassen) AZ, CO, MX, NM
    • intermontana Nelson & Baumann ID, NV, OR, UT
    • manitoba (Claassen) CT, MA, MB, ME, MI, NB, ON, PQ, VT, WI
    • venosa (Banks) ID, MT, OR, WA
    • wanica (Frison) CA, CO, NE, NM, NV, UT, WY
  • Eucapnopsis
    • brevicauda Claassen AB, AK, AZ, BC, CA, CO, ID, MT, NM, NV, OR, SD, UT, WA, WY
  • Isocapnia
    • abbreviata Frison AK, BC, CA, OR, WA
    • agassizi Ricker AB, AK, BC, OR, WA, YT
    • crinita (Needham & Claassen) AB, AK, CO, ID, MT, NM, SK, UT, WY
    • eichlini Zenger & Baumann CA
    • grandis (Banks) AB, AK, BC, CA, ID, MT, OR, UT, WA, WY
    • hyalita Ricker AB, CO, ID, MT, OR, UT, WY
    • integra Hanson AB, AK, BC, ID, MT, SD, WY, YT
    • mogila Ricker CA, OR
    • palousa Zenger & Baumann ID, OR, WA
    • rickeri Zenger & Baumann OR, WA
    • spenceri Ricker AK, BC, CA, OR, WA
    • vedderensis (Ricker) AB, AK, BC, CA, CO, ID, MT, NM, UT, WA
  • Mesocapnia
    • arizonensis (Baumann & Gaufin) AZ, MX, NM
    • autumna (Baumann & Gaufin) AK, BC, OR, WA
    • bakeri (Banks) CA
    • bergi (Ricker) AK
    • bulbosa Nelson & Baumann CA
    • frisoni (Baumann & Gaufin) BJ, CA, CO, MX, NM, TX, UT
    • lapwae (Baumann & Gaufin) BJ, CA, ID, OR
    • oenone (Neave) AB, AK, BC, MT, OR, WA, YK
    • ogotoruka (Jewett) AK
    • porrecta (Jewett) CA, OR, WA
    • projecta (Frison) BC, CA, OR, WA
    • sugluka (Ricker) PQ
    • variabilis (Klapalek) BC
    • werneri (Baumann & Gaufin) AZ, CA, MX, NM
    • yoloensis (Baumann & Gaufin) CA, OR
  • Nemocapnia
    • carolina Banks AL, AR, FL, IL, IN, MS, NC, PQ, SC, VA
  • Paracapnia
    • angulata Hanson AR, CO, CT, DE, IL, KY, LB, MA, MB, MD, ME, MI, MO, NC, NF, NH, NY, OH, OK, PA, PQ, SD, SK, TN, VA, WI, WV, WY
    • boris Stark & Baumann CA
    • disala (Jewett) OR
    • ensicala (Jewett) OR, WA
    • humboldta Baumann & Lee CA
    • opis (Newman) CT, LB, ME, MI, MN, NB, NF, NJ, NY, ON, PQ, WI
  • Utacapnia
    • columbiana (Claassen) AB, AK, BC, CA, MT, OR, YK
    • distincta (Frison) AB, ID, MT, WY
    • imbera (Nebeker & Gaufin) OR, WA
    • labradora (Ricker) LB, PQ
    • lemoniana (Nebeker & Gaufin) CO, ID, NV, SD, UT, WY
    • logana (Nebeker & Gaufin) CO, NM, UT, WY
    • poda (Nebeker & Gaufin) CO, MT, NM, UT, WY
    • sierra (Nebeker & Gaufin) CA
    • tahoensis (Nebeker & Gaufin) CA, NV
    • trava (Nebeker & Gaufin) AB, BC, ID, MB, MT, SK

LEUCTRIDAE
    LEUCTRINAE

  • Calileuctra
    • dobryi Shepard & Baumann CA
    • ephemera Shepard & Baumann CA
  • Despaxia
    • augusta (Banks) AK, BC, CA, ID, MT, OR, WA
  • Leuctra
    • alabama James AL
    • alexanderi Hanson KY, PA, TN, VA, WV
    • alta James AL, IL, IN, KY
    • baddecka Ricker NS
    • biloba Claassen AL, GA, NC, ON, TN, VA
    • carolinensis Claassen MD, NC, TN, VA
    • cottaquilla James AL, FL, MS
    • crossi James AL
    • duplicata Claassen CT, MD, ME, NB, NS, NY, ON, PA, PE, PQ, VA, WV
    • ferruginea (Walker) AL, CT, DE, FL, KY, MA, MD, ME, MI, MN, MS, NB, NC, NF, NS, NY, OH, PA, PQ, SC, SK, VA, WI, WV
    • grandis Banks MA, ME, NB, NC, PA, WV
    • laura Hitchcock NH
    • maria Hanson CT, ME, NH, ON, PA, PQ, VT, WV
    • mitchellensis Hanson NC, VA
    • moha Ricker AL, GA, SC
    • monticola Hanson TN, VA
    • nephophila Hanson NC, TN
    • paleo Poulton & Stewart AR
    • rickeri James AL, IA, IL, IN, KY, MS, OH, WV
    • sibleyi Claassen CT, DE, IL, IN, KY, MA, MD, ME, NB, NC, NY, OH, ON, PA, PQ, TN, VA, WI, WV
    • szczytkoi Stark & Stewart LA
    • tenella Provancher CT, LB, MA, MD, ME, MN, MS, NB, NJ, NS, NY, ON, PA, PQ, WI, WV
    • tenuis (Pictet) AL, AR, CT, DE, IA, IL, IN, KY, MA, MD, ME, MN, MO, MS, NB, NJ, NS, NY, OH, OK, PA, PQ, VA, WI, WV
    • triloba Claassen FL, NC, NY, PQ, SC, VA, WV
    • truncata Claassen CT, ME, NF, NY, PA, PQ, VA, WV
    • variabilis Hanson MA, MD, ME, NH, PA, VA, VT
  • Moselia
    • infuscata (Claassen) BC, CA, NV, OR, WA
  • Paraleuctra
    • alta Baumann & Stark AB
    • divisa (Hitchcock) CA
    • forcipata (Frison) AB, AK, BC, CA, ID, MT, OR, WA
    • jewetti Nebeker & Gaufin CO, NM, MT, UT
    • occidentalis (Banks) AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA, WY
    • projecta (Frison) AK, CO, ID, MT, NM, OR, UT, WA
    • sara (Claassen) AL, CT, DE, KY, MA, MD, ME, NB, NC, NS, NY, OH, ON, PA, PQ, SC, TN, VA, WV
    • vershina Gaufin & Ricker AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, SD, SK, UT, WA, WY
  • Perlomyia
    • collaris Banks BC, CA, ID, OR, WA
    • utahensis Needham & Claassen AB, BC, CA, CO, MT, NM, OR, UT, WA, WY
  • Pomoleuctra
    • andersoni (Harper & Wildman) CA, OR
    • purcellana (Neave) AB, BC, ID, MT, OR, WY
  • Zealeuctra
    • arnoldi Ricker & Ross TX
    • cherokee Stark & Stewart AR, OK
    • claasseni (Frison) AR, IL, IN, KS, KY, MO, OK, TN, TX, WV
    • fraxina Ricker & Ross IL, IN, KY, OH, TN, WV
    • hitei Ricker & Ross TX
    • narfi Ricker & Ross AR, IL, MO, OK, WI
    • stewarti Kondratieff & Zuellig TX
    • talladega Grubbs AL
    • wachita Ricker & Ross AR
    • warreni Ricker & Ross AR, MO, OK

    MEGALEUCTRINAE

  • Megaleuctra
    • complicata Claassen OR, WA
    • flinti Baumann PA, VA, WV
    • kincaidi Frison ID, OR, WA
    • sierra Fields CA
    • stigmata (Banks) AB, BC, MB, MT, WA
    • williamsae Hanson NC, SC, TN, VA

NEMOURIDAE
    AMPHINEMURINAE

  • Amphinemura
    • alabama Baumann AL, KY
    • apache Baumann & Gaufin AZ, MX
    • appalachia Baumann GA, NC, PA, SC, TN, VA
    • banksi Baumann & Gaufin AZ, CO, ID, MT, NM, SD, UT, WY
    • delosa (Ricker) AL, AR, GA, IA, IL, IN, KS, KY, MI, MO, MS, OH, OK, ON, PA, PQ, TN, VA, WI, WV
    • linda (Ricker) AB, AK, BC, IA, LB, MB, MI, NT, ON, PA, PQ, SD, SK, WI
    • mexicana Baumann MX
    • mockfordi (Ricker) TN
    • mogollonica Baumann & Gaufin AZ, NM, UT
    • nigritta (Provancher) AL, AR, CT, DE, FL, GA, IL, IN, KY, LB, MD, ME, MO, MS, NB, NC, NS, NY, OH, ON, PA, PE, PQ, SC, TN, VA, WV
    • puebla Baumann MX
    • reinerti Baumann MX
    • texana Baumann LA, TX
    • varshava (Ricker) IA, IL, IN, KY, WI
    • venusta (Banks) AZ, MX
    • wui (Claassen) CT, DE, GA, KY, MA, MD, ME, NB, NC, NJ, NS, NY, PA, PE, PQ, SC, TN, VA, WV
  • Malenka
    • bifurcata (Claassen) OR, WA
    • biloba (Claassen) BJ, CA
    • californica (Claassen) AB, BC, CA, CO, ID, MB, MT, NM, NV, OR, SK, UT, WA, WY
    • coloradensis (Banks) AZ, CO, NM, SD, UT, WY
    • cornuta (Claassen) BC, CA, OR, WA
    • depressa (Banks) CA, OR
    • flexura (Claassen) AB, CO, ID, MT, NM, OR, WA, WY
    • marionae (Hitchcock) CA
    • perplexa (Frison) OR, WA
    • tina (Ricker) ID, MT, NV, OR, UT, WA
    • wenatchee (Ricker) WA

    NEMOURINAE

  • Lednia
    • tumana (Ricker) MT, WA
  • Nanonemoura
    • wahkeena (Jewett) OR
  • Nemoura
    • arctica Esben-Petersen AB, AK, BC, MB, NT, PQ, SD, WY
    • normani Ricker AK, NT
    • rickeri Jewett AK, MB, SK
    • spiniloba Jewett CA
    • trispinosa Claassen IA, IL, LB, MB, ME, NB, NS, NY, OH, ON, PA, PE, PQ, SD, WI, WY
  • Ostrocerca
    • albidipennis (Walker) CT, MA, MD, ME, NH, NS, NY, ON, PA, PQ, VA, WV
    • complexa (Claassen) CT, MA, ME, NB, NS, NY, ON, PA, PQ, VA, WV
    • dimicki (Frison) BC, OR, WA
    • foersteri (Ricker) BC, OR, WA
    • prolongata (Claassen) DE, ME, NB, NH, NY, PA, PQ, VA, WV
    • truncata (Claassen) CT, IN, KY, MA, MD, ME, NY, OH, ON, PA, PQ, VA, WV
  • Paranemoura
    • claasseni Baumann ME, NB, NH, NS, PE, PQ
    • perfecta (Walker) CT, MA, MD, ME, NB, NC, NH, NS, NY, ON, PA, PE, PQ, TN, VA, VT, WV
  • Podmosta
    • decepta (Frison) AB, AK, BC, CO, ID, MT, OR, UT, WA, WY
    • delicatula (Claassen) AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, SK, UT, WA, WY
    • macdunnoughi (Ricker) LB, ME, NB, NF, NS, PE, PQ
    • obscura (Frison) OR, WA
    • weberi (Ricker) AK, YK
  • Prostoia
    • besametsa (Ricker) AB, BC, CA, CO, ID, MT, NM, NV, OR, SD, UT, WA, WY
    • completa (Walker) AL, AR, DE, IA, IL, IN, KY, MA, ME, MI, MN, MO, MS, NB, NC, NS, OK, ON, PA, PE, PQ, SC, VA, WI, WV
    • hallasi Kondratieff & Kirchner NC, VA
    • similis (Hagen) CT, DE, IN, KY, MA, MD, ME, MI, MN, MO, NY, OH, PA, PQ, SC, VA, WI, WV
  • Shipsa
    • rotunda (Claassen) AB, AK, AL, AR, IA, IL, MB, MD, ME, MI, MN, MS, NB, NT, ON, PQ, SC, SK, VA, WI
  • Soyedina
    • alexandria Grubbs AL, TN
    • calcarea Grubbs KY
    • carolinensis (Claassen) DE, MD, NC, PA, TN, VA, WV
    • interrupta (Claassen) BC, OR, WA
    • kondratieffi Baumann & Grubbs NC
    • merritti Baumann & Grubbs PA
    • nevadensis (Claassen) CA, NV
    • potteri (Baumann & Gaufin) ID, MT
    • producta (Claassen) BC, CA, OR, WA
    • vallicularia (Wu) CT, IA, IL, IN, KY, MA, MD, ME, MI, NS, NY, OH, ON, PA, PQ, TN, VA, WI, WV
    • washingtoni (Claassen) CT, ME, NB, NH, PA, WV
  • Visoka
    • cataractae (Neave) AB, AK, BC, CA, ID, MT, OR, WA
  • Zapada
    • chila (Ricker) NC, TN
    • cinctipes (Banks) AB, AK, BC, CA, CO, ID, MB, MT, NM, NV, SD, SK, UT, WA, WY
    • columbiana (Claassen) AB, AK, BC, CA, ID, MT, OR, UT, WA, WY
    • cordillera (Baumann & Gaufin) CA, ID, MT, OR, WA
    • frigida (Claassen) AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA
    • glacier (Baumann & Gaufin) MT
    • haysi (Ricker) AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA, WY
    • katahdin Baumann & Mingo LB, ME, NH, PQ
    • oregonensis (Claassen) AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, WA, WY, YK

TAENIOPTERYGIDAE
    BRACHYPTERAINAE

  • Bolotoperla
    • rossi (Frison) ME, NC, NH, NY, PA, PQ, VA, WV
  • Doddsia
    • occidentalis (Banks) AK, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA, WY
  • Oemopteryx
    • contorta (Needham & Claassen) CT, KY, MA, MD, ME, NC, NH, PA, TN, VA, WV
    • fosketti (Ricker) AB, CO, MB, MT, ND, SK, UT
    • glacialis (Newport) CT, MI, MN, NY, ON, PQ, WI, WV
    • vanduzeea (Claassen) CA
  • Strophopteryx
    • appalachia Ricker & Ross KY, NC, NY, PA, SC, TN, VA, WV
    • arkansae Ricker & Ross AR, MO, OK
    • cucullata Frison AR, MO, OK, TX
    • fasciata (Burmeister) AL, AR, CT, DE, IA, IL, IN, KS, KY, MB, ME, MI, MN, MO, MS, NC, ND, OH, OK, ON, PA, PQ, SC, VA, WI, WV
    • limata (Frison) NC, TN, VA
  • Taenionema
    • atlanticum Ricker & Ross CT, KY, LB, MA, MD, ME, NB, NC, NF, NH, NS, NY, OH, PA, PQ, SC, TN, VA, VT, WV
    • californicum (Needham & Claassen) CA
    • grinnelli (Banks) CA
    • jacobii Stanger & Baumann AZ, NM
    • jeanae Baumann & Nelson CA
    • jewetti Stanger & Baumann OR, WA
    • kincaidi (Hoppe) AK, BC, CA, OR, WA, YK
    • oregonense (Needham & Claassen) OR, WA
    • pacificum (Banks) AB, AK, AZ, BC, CA, CO, ID, MT, NM, OR, UT, WA, WY, YK
    • pallidum (Banks) AB, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA, WY, YK
    • raynorium (Claassen) CA
    • uinta Stanger & Baumann CO, MT, NV, OR, UT, WY
    • umatilla Stanger & Baumann ID, OR

   TAENIOPTERYGINAE

  • Taeniopteryx
    • burksi Ricker & Ross AL, AR, CO, CT, DE, FL, IA, IL, IN, KS, KY, LA, MD, ME, MI, MN, MO, MS, NC, NE, NY, OH, OK, ON, PA, PQ, SD, TN, TX, VA, WI, WV
    • lita Frison AL, AR, FL, IL, IN, KY, LA, MS, NC, NJ, OK, SC, TX, VA, WV
    • lonicera Ricker & Ross AL, AR, FL, GA, LA, MD, MS, NC, SC, TN, TX, VA
    • maura (Pictet) AL, AR, CT, DE, GA, IN, KY, MA, MD, ME, MI, MN, MS, NC, NY, OH, PA, SC, TN, TX, VA, WV
    • metequi Ricker & Ross AL, AR, IL, IN, KS, KY, MO, NC, NY, OH, OK, ON, PA, VA, WV
    • nelsoni Kondratieff & Kirchner VA
    • nivalis (Fitch) AB, CA, CT, DE, IA, ID, IL, IN, MB, ME, MN, NY, ON, OR, PA, PQ, SK, UT, WI, WA
    • parvula Banks AB, AR, CO, CT, GA, IL, IN, KY, MB, ME, MI, MN, MO, MS, NM, NY, OH, ON, PA, PQ, SC, TN, VA, WI, WV, WY
    • robinae Kondratieff & Kirchner SC
    • starki Stewart & Szczytko TX
    • ugola Ricker & Ross AL, GA, KY, PA, TN, VA, WV

SYSTELLOGNATHA

CHLOROPERLIDAE
    CHLOROPERLINAE

  • Alaskaperla
    • ovibovis (Ricker) AK, NT, YK
  • Alloperla
    • acadiana Harper NB
    • aracoma Harper & Kirchner PA, WV
    • atlantica Baumann AL, CT, GA, MA, MD, ME, MI, MN, NB, NC, NH, NS, NY, OH, ON, PA, PE, PQ, SC, TN, VA, VT
    • banksi Frison ME, MI, NB, NS, NY, ON, PQ, VA
    • biserrata Nelson & Kondratieff MD, PA, VA, WV
    • caddo Poulton & Stewart AR
    • caudata Frison AL, AR, IL, KY, MO, OK
    • chandleri Jewett CA
    • chloris Frison CT, GA, KY, MA, MD, ME, NB, NC, NH, NS, NY, OH, PA, PQ, SC, TN, VA, WV
    • concolor Ricker CT, MA, ME, NB, NF, NH, NS, NY, ON, PA, PQ, VT, WV
    • delicata Frison AB, BC, CA, ID, MT, OR, WA
    • elevata Frison BC, CA, OR, WA
    • fraterna Frison AK, BC, CA, OR, WA
    • furcula Surdick SC
    • hamata Surdick AL, AR, IN, KY, MO
    • idei (Ricker) AL, GA, KY, ME, OH, ON, PQ, VA, WV
    • imbecilla (Say) KY, MD, NY, OH, PA, VA, WV
    • lenati Kondratieff & Kirchner NC
    • leonarda Ricker ME, MI, MN, MO, NB, PQ
    • medveda Ricker AB, BC, ID, MT, WY, YK
    • nanina Banks GA, NC, NY, TN, VA
    • natchez Surdick & Stark MS
    • neglecta Frison NC, TN, VA
    • ouachita Stark & Stewart AR
    • petasata Surdick GA, MA, ME, NB, NC, NF, NH, NS, NY, OH, ON, PA, PQ, SC, TN, VA, VT, WV
    • pilosa Needham & Claassen CO, NM
    • prognoides Surdick & Stark AL
    • roberti Surdick IL Extinct
    • serrata Needham & Claassen AB, AK, BC, ID, MT, WA, WY, YK
    • severa (Hagen) AB, AK, BC, ID, MT, NT, YK
    • stipitata Surdick VA
    • thalia Ricker CO, ID, MT, NV, UT, WY
    • usa Ricker AL, GA, KY, MD, NC, OH, PA, SC, TN, VA, WV
    • voinae Ricker MA, ME, NS, NY, PQ, VT
    • vostoki Ricker ME, NB, NS, NY, PA
  • Bisancora
    • pastina (Jewett) BJ, CA, OR
    • rutriformis Surdick CA
  • Haploperla
    • brevis (Banks) AB, AL, AR, BC, CT, DE, GA, IL, IN, KY, MA, MB, MD, ME, MI, MN, MO, MS, NB, NC, NH, NJ, NS, NY, OH, OK, ON, PA, PE, PQ, SC, SK, TN, VA, VT, WI, WV
    • chilnualna (Ricker) BC, BJ, CA, OR, WA
    • chukcho (Surdick & Stark) MS
    • fleeki Kondratieff, Kirchner & Lenat NC
    • orpha (Frison) ME, MI, MN, NB, ND, ON, PQ, WI
    • parkeri Kondratieff & Kirchner NC
  • Plumiperla
    • diversa (Frison) AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA, WY, YK
    • spinosa (Surdick) CA
  • Rasvena
    • terna (Frison) ME, NC, NH, NY, PA, TN, VA, VT, WV
  • Sasquaperla
    • hoopa Stark & Baumann CA
  • Suwallia
    • amoenacoleus Alexander & Stewart CA
    • autumna (Hoppe) AB, BC, CA, CO, ID, MT, OR, WA, WY
    • dubia (Frison) AB, AK, BC, CA, CO, ID, MT, OR, UT, WA, WY
    • forcipata (Neave) AB, AK, BC, MT, WA, YK
    • lineosa (Banks) AB, AK, BC, CO, ID, MT, OR, SD, SK, UT, WA, WY, YK
    • marginata (Banks) GA, LB, MA, MD, ME, MI, NB, NC, NH, NY, ON, TN, PA, PQ, VA, VT, WI, WV
    • pallidula (Banks) AB, AK, AZ, BC, CA, CO, ID, MB, MT, NM, NV, OR, UT, WA, WY
    • salish Alexander & Stewart MT
    • shepardi Alexander & Stewart CA
    • sierra Baumann & Bottorff CA
    • starki Alexander & Stewart AB, AK, BC, CA, CO, ID, MT, NM, OR, WA, WY
    • sublimis Alexander & Stewart CA
    • wardi Kondratieff & Kirchner CO, MT
  • Sweltsa
    • adamantea Surdick OR, WA
    • albertensis (Needham & Claassen) AB, BC, CO, ID, MT, WY
    • borealis (Banks) AB, AK, BC, CA, CO, ID, MT, NM, NV, OR, SD, UT, WA, WY, YK
    • californica (Jewett) CA
    • coloradensis (Banks) AB, AZ, BC, CA, CO, ID, MT, NM, NV, OR, SD, UT, WA, WY, YK
    • continua (Banks) CA
    • cristata Surdick UT
    • durfeei Kondratieff & Baumann ID, MT
    • exquisita (Frison) AK, BC, OR, WA
    • fidelis (Banks) AB, AK, BC, CA, CO, ID, MT, NV, OR, UT, WA, WY, YK
    • gaufini Baumann ID, UT
    • holstonensis Kondratieff & Kirchner VA
    • hondo Baumann & Jacobi CO, NM
    • lamba (Needham & Claassen) CO, ID, MT, NM, OR, UT, WY
    • lateralis (Banks) CT, GA, MA, MD, ME, NB, NC, NH, NY, OH, PA, PQ, SC, TN, VA, VT, WV
    • mediana (Banks) AL, NC, TN, VA
    • naica (Provancher) LB, ME, NB, NF, NH, NS, NY, PA, PE, PQ, VA, VT, WV
    • occidens (Frison) AK, BC, ID, MT, OR, WA
    • onkos (Ricker) AL, CT, DE, IN, KY, LB, MA, MD, ME, NB, NF, NH, NS, NY, OH, ON, PA, PE, PQ, VA, VT, WV
    • oregonensis (Frison) AK, BC, OR, WA
    • pacifica (Banks) AK, BC, CA, ID, OR, WA
    • palearata Surdick MD, VA, WV
    • pisteri Baumann & Bottorff CA
    • pocahontas Kirchner & Kondratieff MD, WV
    • resima Surdick CA
    • revelstoka (Jewett) AB, AK, BC, MT, OR, WA, WY
    • tamalpa (Ricker) CA
    • townesi (Ricker) CA, NV
    • umbonata Surdick CA
    • urticae (Ricker) NC, VA
    • voshelli Kondratieff & Kirchner GA, NC, VA
    • yurok Stark & Baumann CA
  • Triznaka
    • pintada (Ricker) AZ, CA, CO, ID, NM, NV, OR, SD, UT, WA, WY
    • sheldoni Baumann & Kondratieff CA, ID, NV, UT
    • signata (Banks) AB, AK, BC, CO, ID, MT, NM, OR, SD, SK, UT, WA, WY

    PARAPERLINAE

  • Kathroperla
    • perdita Banks AB, AK, BC, CA, ID, MT, NV, OR, WA
    • takhoma Stark & Surdick CA, WA
  • Paraperla
    • frontalis (Banks) AB, AK, BC, CA, CO, ID, MT, NM, OR, SD, UT, WA, WY, YK
    • wilsoni Ricker AB, BC, CA, ID, MT, OR, WA, YK
  • Utaperla
    • gaspesiana Harper & Roy MD, ME, NH, NY, PA, PQ, WV
    • sopladora Ricker AB, AK, BC, ID, MT, NV, UT, WY, YK

PELTOPERLIDAE
    PELTOPERLINAE

  • Peltoperla
    • arcuata Needham KY, NY, OH, PA, PQ, TN, VA, WV
    • tarteri Stark & Kondratieff VA, WV
  • Sierraperla
    • cora (Needham & Smith) CA, NV, OR
  • Soliperla
    • campanula (Jewett) OR
    • cowlitz Stark & Gustafson WA
    • fenderi (Jewett) WA
    • quadrispinula (Jewett) CA, OR
    • salish Stark & Gustafson ID, MT
    • sierra Stark CA
    • thyra (Needham & Smith) CA, NV
    • tillamook Stark OR
  • Tallaperla
    • anna (Needham & Smith) GA, NC, SC, VA
    • cornelia (Needham & Smith) AL, FL, GA, NC, SC
    • elisa Stark NC, TN
    • laurie (Ricker) AL, GA, SC, TN
    • lobata Stark VA
    • maria (Needham & Smith) AL, CT, DE, GA, KY, MA, MD, ME, NC, NH, NY, PA, SC, TN, VA, WV
    • maiyae Kondratieff, Kirchner & Zuellig NC
  • Viehoperla
    • ada (Needham & Smith) GA, NC, SC, TN
  • Yoraperla
    • brevis (Banks) AB, BC, ID, MT, OR, WY
    • mariana (Ricker) BC, CA, OR, WA
    • nigrisoma (Banks) CA, NV, OR, WA
    • siletz Stark & Nelson BC, CA, OR, WA

PERLIDAE
    ACRONEURIINAE
    ACRONEURINI

  • Acroneuria
    • abnormis (Newman) AB, AL, CO, CT, DE, FL, GA, IA, IL, IN, KS, KY, LA, MA, MB, MD, ME, MI, MN, MS, MT, NB, NC, ND, NE, NF, NM, NY, OH, ON, PA, PQ, SC, SD, SK, TN, UT, VA, WI, WV, WY
    • arenosa (Pictet) AL, CT, DC, DE, FL, GA, LA, MA, MD, ME, MS, NC, NJ, NY, PA, PQ, SC, TX, VA, WV
    • arida (Hagen) GA, NC, NJ, PA, TN
    • carolinensis (Banks) AL, CT, GA, KY, MA, MB, MD, ME, MS, NC, NH, NJ, NY, OH, ON, PA, PQ, SC, TN, VA, WV
    • covelli Grubbs & Stark IN, KY, TN
    • evoluta Klapalek AL, AR, FL, GA, IL, IN, KS, LA, MO, MS, OH, OK, PA, TN, TX
    • filicis Frison AL, AR, GA, IL, IN, KY, MD, MO, OH, PA, SC, TN, VA, WV
    • flinti Stark & Gaufin VA
    • frisoni Stark & Brown AL, AR, IL, IN, KS, KY, MA, MD, MI, MO, NC, OH, OK, ON, PA, TN, VA, WV
    • hitchcocki Kondratieff & Kirchner KY
    • internata (Walker) AR, IL, IN, KY, MI, MN, MO, OK, VA, WI, WV
    • kirchneri Stark & Kondratieff KY, VA, WV
    • kosztarabi Kondratieff & Kirchner VA
    • lycorias (Newman) CT, KY, IA, MB, ME, MI, MN, NB, NC, ND, NS, NY, OH, ON, PA, PQ, SK, TN, VA, WI, WV
    • ozarkensis Poulton & Stewart AR, MO
    • perplexa Frison AL, AR, DC, GA, IL, IN, KY, MO, OH, OK, PA, TN, WV
    • petersi Stark & Gaufin AL, GA, TN
    • yuchi Stark & Kondratieff VA
  • Attaneuria
    • ruralis (Hagen) AL, AR, DC, FL, GA, IA, IL, IN, KS, MB, MD, MN, MO, MS, NC, NE, NY, OH, PA, SC, TN, VA, WI
  • Beloneuria
    • georgiana (Banks) GA, NC
    • jamesae Stark & Szczytko AL
    • stewarti Stark & Szczytko GA, NC, SC, TN
  • Calineuria
    • californica (Banks) AB, BC, CA, ID, MT, OR, WA
  • Doroneuria
    • baumanni Stark & Gaufin BC, CA, NV, OR, WA
    • theodora (Needham & Claassen) AB, BC, ID, MT, WY
  • Eccoptura
    • xanthenes (Newman) AL, CT, DE, FL, GA, KY, MD, MS, NC, NY, OH, PA, SC, TN, VA, WV
  • Hansonoperla
    • appalachia Nelson MA, NH, PA, SC, TN, VA, WV
    • cheaha Kondratieff & Kirchner AL
    • hokolesqua Kondratieff & Kirchner KY, WV
  • Hesperoperla
    • hoguei Baumann & Stark CA
    • pacifica (Banks) AB, AK, AZ, BC, CA, CO, ID, MT, NM, NV, OR, SD, SK, UT, WA, WY
  • Perlesta
    • adena Stark IN, KY, OH, TN
    • baumanni Stark AR, OK
    • bjostadi Kondratieff & Lenat NC
    • bolukta Stark MO, OK
    • browni Stark AR, MO, OK
    • cinctipes (Banks) AR, IA, IL, KS, KY, MO, NE, OH, OK, WV
    • cranshawi Kondratieff & Kirchner VA
    • dakota Kondratieff & Baumann ND, SD
    • decipiens (Walsh) AR, CO, IA, IL, IN, KY, MO, ND, NE, NM, OH, OK, PA, SD, TX, VA, WI, WV, WY
    • etnieri Kondratieff & Kirchner TN
    • frisoni Banks NC, SC, TN, VA, WV
    • fusca Poulton & Stewart AR, MO, OK
    • golconda DeWalt & Stark IA, IL, MO, NE
    • lagoi Stark IA, IL, KY, MS, TN
    • leathermani Kondratieff & Zuellig NC
    • nelsoni Stark NC, PA, SC, TN, WV
    • nitida Banks CT, IN, KY, MA, OH, PA
    • placida (Hagen) AL, DC, FL, GA, LA, MD, ME, MS, NC, PA, TX, VA
    • puttmanni Kondratieff & Kirchner NC, VA
    • roblei Kondratieff & Kirchner NC, VA
    • shawnee Grubbs AL, IL, IN, NC, VA
    • shubuta Stark AR, IA, IL, KY, MI, MO, MS, OK
    • teaysia Kirchner & Kondratieff IL, IN, KY, MD, OH, PA, TN, VA, WV
    • xube Stark & Rhodes IA, IL, ND, NE
  • Perlinella
    • drymo (Newman) AL, AR, CT, DC, FL, GA, IA, IL, IN, KS, KY, LA, MA, MD, ME, MI, MN, MO, MS, NC, NH, NJ, NS, NY, OH, OK, PA, PQ, SC, TN, TX, VA, WI, WV
    • ephyre (Newman) AL, AR, CT, FL, GA, IA, IN, IL, IN, KS, KY, LA, MA, MD, ME, MI, MN, MO, MS, NC, NJ, NY, OH, OK, PA, RI, SC, TN, VA, WI, WV
    • zwicki Kondratieff,et al. AL, FL, MS, SC, NC

    ANACRONEURIINI

  • Anacroneuria
    • litura (Pictet) BE, HO, MX, NI, TX
    • wipukupa Baumann & Olson AZ

    PERLINAE
    NEOPERLINI

  • Neoperla
    • carlsoni Stark & Baumann AL, AR, FL, LA, MO, MS, SC, TX, VA
    • catharae Stark & Baumann AL, AR, IL, IN, KY, MO, OH, OK, PA, TN, VA
    • choctaw Stark & Baumann AR, KY, MO, OK, PA, WV
    • clymene (Newman) AL, FL, GA, IA, IL, IN, KY LA, MS, NC, OK, PA, SC, TX, VA, WV
    • coosa Smith & Stark AL, TN
    • coxi Stark MS
    • falayah Stark & Lentz AR, MO, OK
    • gaufini Stark & Baumann IN, KY, OH
    • harpi Ernst & Stewart AR, IL, MO, OK
    • harrisi Stark & Lentz AL
    • mainensis Banks IL, ME, OH, ON
    • occipitalis (Pictet) AL, IL, IN, KY, ME, MS, NS, NY, OH, ON, PA, SC, TN, VA, WI
    • osage Stark & Lentz AR, IA, IN, MO, OK
    • robisoni Poulton & Stewart AR, IL, MO, MS, OK, PA, TN, WI
    • stewarti Stark & Baumann AL, IL, IN, KY, MA, MD, ME, MN, MS, OH, PA, TN, VA, WI, WV

    PERLINI

  • Agnetina
    • annulipes (Hagen) AL, DC, FL, IN, LA, MD, MS, PA, SC, VA, WV
    • capitata (Pictet) AR, CT, IA, KY, MA, MD, ME, MI, MN, MO, NB, NH, NS, NY, OH, ON, PA, PE, PQ, VA, WI, WV
    • flavescens (Walsh) AL, AR, GA, IL, IN, KY, MD, MN, MO, NC, NY, OH, OK, PA, SC, TN, VA, WV
  • Claassenia
    • sabulosa (Banks) AB, AZ, BC, CA, CO, ID, MB, MT, NM, NT, ON, OR, PQ, SD, SK, UT, WA, WY
  • Paragnetina
    • fumosa (Banks) AL, DC, FL, GA, LA, MS, NC, PA, SC, TX, VA
    • ichusa Stark & Szczytko NC, SC, TN, VA
    • immarginata (Say) CT, DE, GA, KY, MA, ME, NC, NH, NY, PA, PQ, SC, TN, VA, WV
    • kansensis (Banks) AL, AR, FL, GA, IL, IN, KS, LA, MO, MS, NC, SC
    • media (Walker) AR, CT, DE, IA, IL, IN, KY, MB, MD, ME, MI, MN, MO, NB, NH, NY, OH, ON, PA, PQ, SK, VA, WI, WV

PERLODIDAE
    ISOPERLINAE

  • Calliperla
    • luctuosa (Banks) CA, OR, WA
  • Cascadoperla
    • trictura (Hoppe) BC, CA, ID, MT, OR, WA
  • Clioperla
    • clio (Newman) AL, AR, CT, DE, FL, GA, IA, IL, IN, KY, MA, MD, MI, MO, MS, NC, OH, OK, ON, PA, SC, TN, VA, WI, WV
  • Cosumnoperla
    • hypocrena Szczytko & Bottorff CA
    • sequoia Bottorff CA
  • Isoperla
    • acula Jewett CA
    • adunca Jewett CA
    • baumanni Szczytko & Stewart CA
    • bellona Banks GA, NC
    • bifurcata Szczytko & Stewart CA, ID, OR, WA
    • bilineata (Say) AL, CO, CT, IA, IL, IN, KS, KY, MA, MB, ME, MI, MN, MO, MS, NB, NC, ND, NE, NF, NY, OH, ON, PA, PQ, SD, SK, VA, WI, WV
    • burksi Frison AR, IL, IN, KY, MD, MO, NC, OH, OK, SC, WV
    • conspicua Frison IL Extinct
    • cotta Ricker MA, ME, MI, NB, NS, ON, PE, PQ, SC, WI, WV
    • coushatta Szczytko & Stewart AR, DE, LA, MS, OK, TX
    • davisi James AL, SC
    • decepta Frison IL, IN, KY, MO, OH, ON
    • decolorata (Walker) AK, BC, MB, NT, SK, YK
    • denningi Jewett CA
    • dicala Frison AL, AR, CT, FL, IA, IN, KY, MA, MB, MD, ME, MI, MN, MO, MS, NB, NC, ON, PA, PQ, SC, TN, VA, WI, WV
    • distincta Nelson NC, SC, TN
    • emarginata Harden & Mickel MN
    • francesca Harper CT, MA, ME, NY, PA, PQ
    • frisoni Illies CT, DE, IN, MB, ME, MI, MN, NB, NC, NS, ON, PA, PE, PQ, WI
    • fulva Claassen AB, AZ, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA, WY
    • fusca Needham & Claassen AB, BC, ID, MT, OR, WA, WY, YK
    • gibbsae Harper CT, MD, NY, PQ, WV
    • gravitans (Needham & Claassen) OR, WA
    • holochlora (Klapalek) CT, DE, GA, KY, MA, MD, ME, NC, NS, NY, PA, PQ, TN, VA, WV
    • irregularis (Klapalek) TX
    • jewetti Szczytko & Stewart CO, NM, TX
    • karuk Baumann & Lee CA
    • katmaiensis Szczytko & Stewart AK
    • lata Frison MA, ME, MI, MN, NB, NC, NS, ON, PA, PQ, TN, VA, WI, WV
    • laucki Baumann & Lee CA
    • longiseta Banks AB, BC, CO, IA, ID, IL, KS, MB, MN, MO, MT, ND, NM, PQ, SD, SK, UT, WY
    • major Nelson & Kondratieff VA
    • marlynia (Needham & Claassen) CO, IA, IL, IN, KS, KY, MB, ME, MI, MN, NB, NE, NH, NJ, NS, OK, ON, PA, PQ, SC, SK, VA, WI, WV
    • marmorata Needham & Claassen CA, NV, OR, WA
    • maxana Harden & Mickel MN
    • miwok Bottorff & Szczytko CA
    • mohri Frison AR, IL, KS, LA, MO, OK
    • montana (Banks) CT, DE, ME, MN, NH, NS, NY, ON, PA, PQ
    • mormona Banks AZ, BC, BJ, CA, CO, ID, MT, NM, OR, UT, WA, WY
    • muir Szczytko & Stewart OR
    • namata Frison AR, IN, KY, ME, MO, OH, PA, VA, WV
    • nana (Walsh) IL, IN, KY, MI, OH, ON, PA, PQ, WI
    • orata Frison CT, FL, KY, ME, MN, NB, NC, NH, NS, NY, OH, PA, PQ, SC, TN, VA, VT, WV
    • ouachita Stark & Stewart AR, MO, OK
    • petersoni Needham & Christenson AB, AK, BC, CO, ID, MT, SK, UT, WA, WY, YK
    • phalerata (Smith) CO, ID, NM, OR, SD, UT, WY
    • pinta Frison AB, BC, BJ, CA, CO, ID, MT, OR, UT, WA, WY
    • quinquepunctata (Banks) AB, BC, BJ, CA, CO, ID, KS, MT, NE, NM, NV, OR, SD, SK, UT, WY
    • rainieri Jewett OR, WA
    • richardsoni Frison CT, IA, IL, KY, MN, MO, PA, WI, WV
    • roguensis Szczytko & Stewart CA, OR
    • sagittata Szczytko & Stewart TX
    • signata (Banks) AR, CT, IA, MB, ME, MI, MN, MO, NB, NS, NY, OH, OK, PA, PQ, VA, WI, WV
    • similis (Hagen) CT, DE, KY, MA, MD, ME, NC, NH, NY, PA, PQ, TN, VA, WV
    • slossonae (Banks) CT, IA, ME, MI, MN, NB, NC, NH, NS, NY, PA, PE, PQ, VA, WI, WV
    • sobria (Hagen) AB, AK, AZ, BC, CA, CO, ID, MT, NM, NV, OR, UT, WA, WY, YK
    • sordida Banks AB, AK, BC, CA, ID, MT, OR, WA
    • szczytkoi Poulton & Stewart AR
    • tilasqua Szczytko & Stewart OR, WA
    • transmarina (Newman) AB, BC, CT, DE, IA, KY, LB, MB, ME, MI, MN, NB, NC, NF, NJ, NS, NY, ON, PA, PE, PQ, SD, SK, VA, WI, WV, WY

    PERLODINAE
    ARCYNOPTERYGINI

  • Arcynopteryx
    • compacta (McLachlan) AB, AK, BC, CO, ME, MI, MT, NH, NT, NY, SK, WY
  • Frisonia
    • picticeps (Hanson) BC, CA, NV, OR, WA
  • Megarcys
    • irregularis (Banks) BC, WA
    • signata (Hagen) AK, BC, CO, ID, MT, NM, NV, UT, WY
    • subtruncata Hanson BC, ID, MT, OR, WA
    • watertoni (Ricker) AB, BC, ID, MT
    • yosemite (Needham & Claassen) CA, WA
  • Oroperla
    • barbara Needham CA
  • Perlinodes
    • aurea (Smith) AB, CA, ID, MT, OR, WA, WY
  • Salmoperla
    • sylvanica Baumann & Lauck CA
  • Setvena
    • bradleyi (Smith) AB, BC, ID, MT
    • tibialis (Banks) BC, OR, WA
    • wahkeena Stewart & Stanger OR
  • Skwala
    • americana (Klapalek) AZ, BC, CA, CO, ID, MB, MT, NM, NV, OR, SD, SK, UT, WA, WY
    • curvata (Hanson) BC, CA, ID, MT, OR, WA, WY

    DIPLOPERLINI

  • Baumannella
    • alameda (Needham & Claassen) CA
  • Cultus
    • aestivalis (Needham & Claassen) AZ, BC, CO, ID, MT, NM, NT, UT, WY, YK
    • decisus decisus (Walker) MI, NB, NY, ON, PA
    • decisus isolatus (Banks) GA, NC, VA
    • pilatus (Frison) BC, CA, ID, MT, OR, WA
    • tostonus (Ricker) BC, CA, ID, MT, OR, WA, WY
    • verticalis (Banks) NC, NH, PA, PQ, TN, VA, WV
  • Diploperla
    • duplicata (Banks) AL, DE, GA, MD, MS, NC, PA, SC, TN, VA, WV
    • kanawholensis Kirchner & Kondratieff KY, VA, WV
    • morgani Kondratieff & Voshell NC, VA
    • robusta Stark & Gaufin CT, IN, KY, OH, PA, TN, VA, WV
  • Kogotus
    • modestus (Banks) BC, CO, ID, MT, NM, UT, WY
    • nonus (Needham & Claassen) AK, BC, CA, ID, MT, OR, WA, WY
  • Osobenus
    • yakimae (Hoppe) BC, CA, OR, WA
  • Pictetiella
    • expansa (Banks) CO, ID, MT, UT, WY
    • lechleitneri Stark & Kondratieff WA
  • Remenus
    • bilobatus (Needham & Claassen) AL, CT, DE, GA, KY, MD, NC, NY, PA, SC, TN, VA, WV
    • duffieldi Nelson & Kondratieff GA
    • kirchneri Kondratieff & Nelson VA, WV
  • Rickera
    • sorpta (Needham & Claassen) CA, NV, OR, WA

    PERLODINI

  • Chernokrilus
    • misnomus (Claassen) CA, OR
  • Diura
    • bicaudata (Linnaeus) AK, MB, NT, SK, YK
    • knowltoni (Frison) AB, BC, CA, CO, ID, MT, NM, NV, OR, SK, UT, WY, YK
    • nanseni (Kempny) NH, PQ
  • Helopicus
    • bogaloosa Stark & Ray AL, FL, GA, LA, MS, NC, SC
    • nalatus (Frison) AR, IN, KS, MI, MO, OK
    • subvarians (Banks) AL, CT, FL, KY, ME, NC, ON, PA, PQ, SC, TN, VA, WV
  • Hydroperla
    • crosbyi (Needham & Claassen) AR, IL, IN, KS, MO, OK, TX
    • fugitans (Needham & Claassen) AR, IA, IL, IN, KS, MO, TN, TX
    • phormidia Ray & Stark FL, NC, SC
    • rickeri (Stark) TN
  • Isogenoides
    • colubrinus (Hagen) AB, AK, AZ, BC, CA, CO, ID, MB, MT, NM, NT, OR, SK, UT, WA, WY, YK
    • doratus (Frison) IA, BC, MB, MN, MI, PQ
    • elongatus (Hagen) AB, AZ, BC, CO, ID, MB, MT, NM, SD, UT, WA, WY
    • frontalis (Newman) MB, ME, MI, MN, NF, NY, ON, PQ, SK, WI
    • hansoni (Ricker) CT, MA, MD, ME, NB, NC, NS, NY, PA, PQ, VA, WV
    • olivaceus (Walker) MI, MN, ON, PA, PQ, WI
    • varians (Walsh) IA, IL, IN, KS, MI, MN, MS, NC, SC, TN, VA
    • zionensis Hanson AK, AZ, BC, CO, NM, NT, UT
  • Malirekus
    • hastatus (Banks) GA, KY, NC, SC, TN, VA, WV
    • iroquois Stark & Szczytko MD, NB, NH, NY, PA, PQ, VT
  • Oconoperla
    • innubila (Needham & Claassen) NC, SC, TN
  • Susulus
    • venustus (Jewett) CA
  • Yugus
    • arinus (Frison) GA, NC, SC, TN, VA
    • bulbosus (Frison) GA, NC, TN
    • kirchneri Nelson KY, MD, PA, VA, WV
    • kondratieffi Nelson NC, VA

PTERONARCYIDAE
    PTERONARCYINAE
    PTERONARCELLINI

  • Pteronarcella
    • badia (Hagen) AB, AK, AZ, BC, CO, ID, MT, NM, NV, OR, SK, UT, WY
    • regularis (Hagen) AB, AK, CA, NV, OR, WA

    PTERONARCYINI

  • Pteronarcys
    • biloba Newman AL, CT, GA, KY, MA, MD, ME, NB, NC, NS, NY, PA, PE, PQ, VA, WV
    • californica Newport AK, AZ, BC, CA, CO, ID, MT, MX, NM, OR, UT, WA, WY
    • comstocki Smith KY, ME, NB, NY, PA, VA, WV
    • dorsata (Say) AB, AK, AL, BC, FL, KS, KY, LA, LB, MB, MD, ME, MI, MN, MS, MT, NB, NC, NY, OH, PA, PQ, SC, SK, TN, VA, WI, WV, WY
    • pictetii Hagen AR, CO, CT, IA, IL, IN, KS, KY, MB, MI, MN, MO, ND, NE, OH, PA, SD, TN, WI
    • princeps Banks BC, CA, ID, NV, OR, UT, WA
    • proteus Newman KY, MB, MD, ME, NC, NH, NY, PA, PQ, VA, WV
    • scotti Ricker GA, NC, SC, TN, VA

This list contains 674 valid North American stonefly species.
Contact R. Edward DeWalt, edewalt@inhs.uiuc.edu, to submit new valid names and published state and province records. Citation or pdf of article must accompany request as proof of claim.



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